Ray Romano

Manny (voice)

John Leguizamo

Sid (voice)

Denis Leary

Diego (voice)

Adam Devine

Julian (voice)

Jesse Tyler Ferguson

Shangri Llama (voice)

Max Greenfield

Roger (voice)

Jessie J

Brooke (voice)

Nick Offerman

Gavin (voice)

Keke Palmer

Peaches (voice)

Josh Peck

Eddie (voice)

Simon Pegg

Buck / Pythagorus Buck / Robo Buck (voice)

Seann William Scott

Crash (voice)

Wanda Sykes

Granny (voice)

Jennifer Lopez

Shira (voice)

Queen Latifah

Ellie (voice)

Stephanie Beatriz

Gertie (voice)

Neil deGrasse Tyson

Neil deBuck Weasel (voice)

Lilly Singh

Bubbles and Misty (voice)

Michael Strahan

Teddy (voice)

Melissa Rauch

Francine (voice)

Chris Wedge

Scrat (voice)

Robert Cardone

Freaky Bobby (voice)

Byron Thames

Additional Voices (voice)

Rif Hutton

Additional Voices (voice)

Chris Campbell

Additional Voices (voice)

Holly Dorff

Additional Voices (voice)

Moosie Drier

Additional Voices (voice)

Jason Fricchione

Additional Voices (voice)

Jackie Gonneau

Additional Voices (voice)

Hope Levy

Additional Voices (voice)

Terence Mathews

Additional Voices (voice)

Edie Mirman

Additional Voices (voice)

Juan Pope

Additional Voices (voice)

Ruth Zalduondo

Additional Voices (voice)

David Zyler

Additional Voices (voice)

Ashley Peldon

Additional Voices (voice)

Carlos Ponce

Mariachi Beaver (voice)

Michael Thurmeier

Gravedigger Beaver / Party Molehog (voice)

Randy Thom

Hyrax Minister (voice)

Jorge Lucas

Grandpa Molehog (voice)

James Palumbo

Panicked Start (voice)